[RodB]

Now you're on track! I was just writing a response and noticed you figured it out. You have to be very careful with wiring this section. On the schematic there are several wires coming off the candohm running to the grid circuits of other tubes. These are negative bias voltages that control the current flow through those tubes. The candohm sets up the voltages. When you grounded both ends of the candohm these voltages were gone. Note that when you tell us you've wired in a part it helps to use the schematic designated number to make it easier for us to follow along.